Biology:Trogulidae

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Short description: Family of harvestmen/daddy longlegs

Trogulidae
Temporal range: Palaeogene–present
Trogulus tricarinatus.jpg
Trogulus tricarinatus
Scientific classification e
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Opiliones
Suborder: Dyspnoi
Superfamily: Troguloidea
Family: Trogulidae
Sundevall, 1833
Genera
  • Amphitrogulus
  • Anarthrotarsus
  • Anelasmocephalus
  • Calathocratus
  • Kofiniotis
  • Trogulus
Diversity
6 genera, 65 species
Distribution.trogulidae.1.png
Approximate natural range of Trogulidae

Trogulidae is a family of harvestmen comprising 65 extant species (plus 1 extinct) in five genera.

Members of this species have short legs and live in soil. They have dirt attached to their bodies, to escape predators. Their body length ranges from 2 to 22 mm. In most genera the body is somewhat flattened and leathery. Adults have a small hood which hides their short chelicerae and pedipalps.[1]

Distribution

Members of this family occur in western and southern Europe, up to western North Africa and the Levant, the Caucasus and northern Iran. Trogulus tricarinatus, a predator of terrestrial snails, has been introduced to eastern North America.[1]

Name

The derivation of the name of the type genus, Trogulus, is not fully understood. The describer Latreille wrote that he named it because it looks like a monkshood. Perrier (1929) however derived the name from Ancient Greek trogein "gnawing", because of the rough, "gnawed-upon" appearance.[1]

Species

Main page: Biology:List of Dyspnoi species

The known genera and species in Trogulidae as of 2023 are:[2][3][4]

  • Anarthrotarsus Šilhavý, 1967
  • Anarthrotarsus martensi Šilhavý, 1967 – Greece
  • Anarthrotarsus trichasi Kontos & Martens, 2022 – Greece
  • Anelasmocephalus balearicus Martens & Chemini, 1988 – Spain (Baleares)
  • Anelasmocephalus brignolii Martens & Chemini, 1988 – Italy (Sardinia)
  • Anelasmocephalus calcaneatus Martens & Chemini, 1988 – Italy (Sicily)
  • Anelasmocephalus cambridgei (Westwood, 1874) – UK to Balkans (Europe)
  • Anelasmocephalus cazorla Prieto & Las Heras, 2020 – Spain
  • Anelasmocephalus crassipes (H. Lucas, 1846) – Algeria, Tunisia
  • Anelasmocephalus gadirrif Prieto & Las Heras, 2020 – Spain, Morocco
  • Anelasmocephalus hadzii Martens, 1978 – Austria, Italy, Serbia, etc Balkans
  • Anelasmocephalus lycosinus (Sørensen, 1873) – Italy
  • Anelasmocephalus ortunioi Prieto & Las Heras, 2020 – Spain
  • Anelasmocephalus osellai Martens & Chemini, 1988 – Italy
  • Anelasmocephalus pusillus Simon, 1879 – Italy, France (Corsica)
  • Anelasmocephalus pyrenaicus Martens, 1978 – Spain
  • Anelasmocephalus rufitarsis Simon, 1879 – Italy, France, Switzerland
  • Anelasmocephalus tenuiglandis Martens & Chemini, 1988 – France
  • Anelasmocephalus tuscus Martens & Chemini, 1988 – Italy
  • Calathocratus Simon, 1879
  • Calathocratus africanus (H. Lucas, 1846) – Algeria, Tunisia, Italy
  • Calathocratus beieri Gruber, 1968 – Turkey
  • Calathocratus caucasicus (Šilhavý, 1966) – Turkey, Azerbaijan, Russia, Georgia, Iran.
  • Calathocratus hirsutus Snegovaya, 2011 – Russia
  • Calathocratus intermedius Roewer, 1940 – Greece (Crete)
  • Calathocratus kyrghyzicus (Chemeris, 2013) – Kyrgyzstan
  • Calathocratus minutus Snegovaya, 2011 – Russia
  • Calathocratus rhodiensis (Gruber, 1963) – Greece (Rhodes)
  • Calathocratus singularis (Roewer, 1940) – Greece, Bulgaria, Turkey
  • Calathocratus sinuosus (Sørensen, 1873) – Italy
  • Kofiniotis Roewer, 1940
  • Kofiniotis creticus Roewer, 1940 – Greece (Crete)
  • Trogulus Latreille, 1802
  • Trogulus aquaticus Simon, 1879 – France (Corsica)
  • Trogulus balearicus Schönhofer & Martens, 2008 – Spain (Baleares)
  • Trogulus banaticus Avram, 1971 – Romania, North Macedonia, Serbia, etc.
  • Trogulus cisalpinus Chemini & Martens, 1988 – Italy
  • Trogulus closanicus Avram, 1971 – Romania, Germany, Bulgaria etc
  • Trogulus coreiformis C. L. Koch, in Hahn & Koch, 1839 – (Unknown locality) [Nomen dubium per Schönhofer 2013]
  • Trogulus coriziformis C. L. Koch, in Hahn & Koch, 1839  – Italy (contra reports of widespread)
  • Trogulus cristatus Simon, 1879 – France, Italy
  • Trogulus falcipenis Komposch, 1999 – Austria, Italy, Croatia, Slovenia
  • Trogulus graecus Dahl, 1903 – Greece, Albania, etc (Balkans)
  • Trogulus gypseus Simon, 1879 – Israel, Egypt, Turkey, Greece.
  • Trogulus hirtus Dahl, 1903 – Bosnia and Herzegovina, Croatia
  • Trogulus huberi Schönhofer & Martens, 2008 – Portugal
  • Trogulus karamanorum Schönhofer & Martens, 2009 – Macedonia, Albania etc (Balkans)
  • Trogulus lusitanicus Giltay, 1932  – Portugal
  • Trogulus lygaeiformis C. L. Koch, in Hahn & Koch, 1839  – Greece [Nomen dubium per Schönhofer 2013]
  • Trogulus martensi Chemini, 1983  – Italy, France, etc
  • Trogulus megaligrava Schönhofer et al., 2013 – Greece (Corfu)
  • Trogulus melitensis Schönhofer & Martens, 2009 – Croatia
  • Trogulus nepaeformis (Scopoli, 1763) – Italy, France, etc (Southern Europe)
  • Trogulus oltenicus Avram, 1971 – Romania
  • Trogulus ozimeci Schönhofer et al., 2013 – Croatia
  • Trogulus pharensis Schönhofer & Martens, 2009 – Croatia
  • Trogulus prietoi Schönhofer & Martens, 2008 – Spain
  • Trogulus pulverulentus C. L. Koch, 1856 – Spain [Nomen dubium per Schönhofer 2013]
  • Trogulus pyrenaicus Schönhofer & Martens, 2008 – Spain
  • Trogulus rossicus Šilhavý, 1968 – Russia, Azerbaijan, Georgia
  • Trogulus setosissmus Roewer, 1940 – Croatia
  • Trogulus squamatus C. L. Koch, in Hahn & Koch, 1839 – Montenegro, Croatia (Dalmatia)
  • Trogulus templetonii Westwood, 1833 – Spain [Nomen dubium per Schönhofer 2013]
  • Trogulus tenuitarsus Schönhofer et al., 2013 – Macedonia
  • Trogulus thaleri Schönhofer & Martens, 2009 – Croatia
  • Trogulus tingiformis C. L. Koch, in Hahn & Koch, 1839 – Germany, etc (Central Europe)
  • Trogulus torosus Simon, 1885 – Montenegro, Bosnia and Herzegovina, Croatia (Dalmatia)
  • Trogulus tricarinatus (Linnaeus, 1758) – Germany, etc (Widespread Western, Central Europe)
  • Trogulus uncinatus Gruber, 1973 – Turkey

See also extinct:

  • Trogulus longipes Haupt, 1956 (fossil: Eocene)

Plus see unknown:

  • Amphitrogulus Gourret, 1887
  • Amphitrogulus sternalis Gourret, 1887

References

  1. 1.0 1.1 1.2 Gruber, Jürgen (2007): Trogulidae Sundevall, 1833. In: Pinto-da-Rocha et al. 2007: 157ff
  2. "A taxonomic catalogue of the Dyspnoi Hansen and Sørensen, 1904 (Arachnida: Opiliones)" (in en). https://www.researchgate.net/publication/240310711. 
  3. Chemeris, A.N. (2013) Two new harvestman species (Arachnida: Opiliones) from the collection of Siberian Zoological Museum. Arthropoda Selecta
  4. "Trogulidae". Species. GBIF. http://www.gbif.org/species/3253446. 

Further reading

  • Joel Hallan's Biology Catalog: Trogulidae
  • Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds.) (2007): Harvestmen - The Biology of Opiliones. Harvard University Press ISBN:0-674-02343-9

Wikidata ☰ Q2312673 entry