Biology:List of peramelemorphs

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Short description: Species in mammal order Peramelemorphia

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brown bandicoot
Eastern barred bandicoot (Perameles gunni)
brown bilby
Greater bilby (Macrotis lagotis)

Peramelemorphia is an order of Australian marsupial mammals. Members of this order are called peramelemorphs and include bandicoots and bilbies. They are found in Australia and New Guinea, generally in forests, shrublands, grasslands, and savannas, though some species are found in inland wetlands and deserts. They range in size from the Arfak pygmy bandicoot, at 14 cm (6 in) plus a 11 cm (4 in) tail, to the giant bandicoot, at 56 cm (22 in) plus a 34 cm (13 in) tail. Peramelemorphs primarily eat insects and fruit, as well as other invertebrates and small vertebrates. Most peramelemorphs do not have population estimates, but the ones that do range from 1,000 to 100,000 mature individuals. The giant bandicoot, David's echymipera, and Seram bandicoot are endangered. The desert bandicoot, lesser bilby, and pig-footed bandicoot were driven to extinction in the mid-1900s.

The nineteen extant species of Peramelemorphia are divided into two families: Peramelidae, containing eighteen species divided between three genera in the subfamily Echymiperinae, two genera in the subfamily Peramelinae, and a single genus in the subfamily Peroryctinae; and Thylacomyidae, containing one extant species in a single genus. Additionally, Peramelemorphia includes the extinct family Chaeropodidae, containing a single species. Dozens of extinct, prehistoric Peramelemorphia species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[1]

Conventions

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the peramelemorph's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "Extinct".

Classification

Peramelemorphia[2]  
ChaeropodidaeExtinct  

ChaeropusExtinct

Peramelidae  
Peramelinae  

Isoodon

Perameles

Echymiperinae  

Echymipera

Microperoryctes

Rhynchomeles

Peroryctinae  

Peroryctes

Thylacomyidae  

Macrotis

The order Peramelemorphia consists of nineteen extant species in two extant families, Peramelidae and Thylacomyidae. Peramelidae is divided into three subfamilies: Echymiperinae, containing ten species in three genera; Peramelinae, containing six species in two genera; and Peroryctinae, containing a single species. Thylacomyidae consists of one extant species. Additionally, Peramelemorphia includes the extinct family Chaeropodidae, containing a single species. Three species have been driven to extinction in recent times, all in the mid-1900s: the desert bandicoot in Peramelinae, the lesser bilby in Thylacomyidae, and the pig-footed bandicoot in Chaeropodidae. Many of these species are further subdivided into subspecies. This does not include hybrid species or extinct prehistoric species.

Family ChaeropodidaeExtinct

  • Genus ChaeropusExtinct (pig-footed bandicoots): one species (one extinct)

Family Peramelidae

  • Subfamily Echymiperinae
  • Subfamily Peramelinae
    • Genus Isoodon (short-nosed bandicoots): three species
    • Genus Perameles (long-nosed bandicoots): four species (one extinct)
  • Subfamily Peroryctinae
    • Genus Peroryctes (New Guinean long-nosed bandicoots): two species

Family Thylacomyidae

  • Genus Macrotis (bilbies): two species (one extinct)

Peramelemorphs

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[3]

Chaeropodidae

Genus ChaeropusExtinctOgilby, 1838 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Pig-footed bandicootExtinct

Brown bandicoot

C. ecaudatus[lower-alpha 1]
(Ogilby, 1838)
Central and western Australia
Map of range
Size: 23–26 cm (9–10 in) long, plus 10–14 cm (4–6 in) tail[4]

Habitat: Forest, savanna, shrubland, and grassland[5]

Diet: Believed to be omnivorous[4]
 EX 


0 Population steady[5]

Peramelidae

Subfamily Echymiperinae

Genus EchymiperaLesson, 1842 – five species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Clara's echymipera E. clara
Stein, 1932
Northern New Guinea
Map of range
Size: 20–50 cm (8–20 in) long, plus 5–13 cm (2–5 in) tail[6]

Habitat: Forest[7]

Diet: Omnivorous, primarily fruit[6]
 LC 


Unknown Template:Population change unknown[7]

Common echymipera E. kalubu
(J. B. Fischer, 1829)

New Guinea
Map of range
Size: 22–38 cm (9–15 in) long, plus 8 cm (3 in) tail[8]

Habitat: Inland wetlands and forest[9]

Diet: Invertebrates and fruit[8]
 LC 


Unknown Population steady[9]

David's echymipera E. davidi
Flannery, 1990
Kiriwina island east of New Guinea
Map of range
Size: 20–50 cm (8–20 in) long, plus 5–13 cm (2–5 in) tail[6]

Habitat: Forest[10]

Diet: Omnivorous[6]
 EN 


Unknown Template:Population change unknown[10]

Long-nosed echymipera E. rufescens
(Peters, Doria, 1875)

New Guinea and northeastern Australia
Map of range
Size: 20–50 cm (8–20 in) long, plus 5–13 cm (2–5 in) tail[11]

Habitat: Forest, shrubland, and grassland[12]

Diet: Fruit, seeds, invertebrates, fungi, and plants[11]
 LC 


Unknown Population declining[12]

Menzies' echymipera E. echinista
Menzies, 1990
Southern New Guinea
Map of range
Size: 20–50 cm (8–20 in) long, plus 5–13 cm (2–5 in) tail[6]

Habitat: Savanna and forest[13]

Diet: Omnivorous[6]
 DD 


Unknown Template:Population change unknown[13]

Genus MicroperoryctesStein, 1932 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Arfak pygmy bandicoot M. aplini
Helgen, Flannery, 2004
Western New Guinea
Map of range
Size: 14–16 cm (6 in) long, plus 11–12 cm (4–5 in) tail[14]

Habitat: Forest[15]

Diet: Insects as well as fruit[16]
 VU 


Unknown Template:Population change unknown[15]

Mouse bandicoot M. murina
Stein, 1932
Western New Guinea
Map of range
Size: 15–18 cm (6–7 in) long, plus 10–11 cm (4 in) tail[17]

Habitat: Forest[18]

Diet: Insects as well as fruit[16]
 VU 


Unknown Template:Population change unknown[18]

Papuan bandicoot M. papuensis
(Laurie, 1952)
Eastern New Guinea
Map of range
Size: 17–20 cm (7–8 in) long, plus 13–16 cm (5–6 in) tail[17]

Habitat: Forest[19]

Diet: Insects as well as fruit[16]
 LC 


Unknown Template:Population change unknown[19]

Striped bandicoot

Drawing of brown bandicoot

M. longicauda
(Peters, Doria, 1876)

New Guinea
Map of range
Size: 23–31 cm (9–12 in) long, plus 14–26 cm (6–10 in) tail[17]

Habitat: Forest and grassland[20]

Diet: Insects as well as fruit[16]
 LC 


Unknown Template:Population change unknown[20]

Genus RhynchomelesThomas, 1920 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Seram bandicoot R. prattorum
Thomas, 1920
Seram Island west of New Guinea
Map of range
Size: 24–33 cm (9–13 in) long, plus 10–13 cm (4–5 in) tail[21]

Habitat: Forest[22]

Diet: Unknown[21]
 EN 


Unknown Template:Population change unknown[22]

Subfamily Peramelinae

Genus IsoodonDesmarest, 1817 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Golden bandicoot

Brown bandicoot

I. auratus
(Ramsay, 1887)

Northwestern Australia
Map of range
Size: 19–30 cm (7–12 in) long, plus 8–13 cm (3–5 in) tail[23]

Habitat: Savanna, shrubland, and grassland[24]

Diet: Insects and arachnids, as well as turtle eggs, small reptiles, and plant material[25]
 VU 


30,000 Population declining[24]

Northern brown bandicoot

Brown bandicoot

I. macrourus
(Gould, 1842)

Northern and eastern Australia and southern New Guinea (former in black)
Map of range
Size: 30–47 cm (12–19 in) long, plus 8–22 cm (3–9 in) tail[26]

Habitat: Forest, savanna, shrubland, and grassland[27]

Diet: Insects, as well as lizards, birds, plant matter, and fungi[26]
 LC 


Unknown Population steady[27]

Southern brown bandicoot

Brown bandicoot

I. obesulus
(Shaw, 1797)

Southwestern and southeastern Australia
Map of range
Size: 28–36 cm (11–14 in) long, plus 9–14 cm (4–6 in) tail[28]

Habitat: Forest, shrubland, grassland, and inland wetlands[29]

Diet: Plants, fungi, and invertebrates[30]
 LC 


10,000–100,000 Population declining[29]

Genus PeramelesGeoffroy, 1803 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Desert bandicootExtinct

Brown bandicoot

P. eremiana
Spencer, 1897
Central Australia Size: 20–43 cm (8–17 in) long, plus 7–17 cm (3–7 in) tail[31]

Habitat: Shrubland, grassland, and desert[32]

Diet: Believed to be insects, as well as worms, snails, mice, lizards, and plants[31]
 EX 


0 Population steady[32]

Eastern barred bandicoot

Brown bandicoot

P. gunnii
Gray, 1838
Southern Australia
Map of range
Size: 20–43 cm (8–17 in) long, plus 7–17 cm (3–7 in) tail[31]

Habitat: Grassland[33]

Diet: Insects and arachnids, as well as small vertebrates and plants[34]
 VU 


10,000–100,000 Population declining[33]

Long-nosed bandicoot

Brown bandicoot

P. nasuta
Geoffroy, 1804
Eastern Australia
Map of range
Size: 20–43 cm (8–17 in) long, plus 7–17 cm (3–7 in) tail[31]

Habitat: Forest, shrubland, and grassland[35]

Diet: Insects, arachnids, small vertebrates, plant roots, and fungi[36]
 LC 


Unknown Template:Population change unknown[35]

Western barred bandicoot

Brown bandicoot

P. bougainville
Quoy, Gaimard, 1824
Scattered western and central Australia
Map of range
Size: 20–43 cm (8–17 in) long, plus 7–17 cm (3–7 in) tail[31]

Habitat: Shrubland and grassland[37]

Diet: Invertebrates, plants, roots, herbs, seeds, berries, and fungi, as well as small vertebrates[38]
 VU 


1,000–5,000 Population steady[37]

Subfamily Peroryctinae

Genus PeroryctesThomas, 1906 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Giant bandicoot P. broadbenti
(Ramsay, 1879)
Eastern New Guinea
Map of range
Size: 39–56 cm (15–22 in) long, plus 11–34 cm (4–13 in) tail[39]

Habitat: Forest[40]

Diet: Believed to be omnivorous[40]
 EN 


Unknown Population declining[40]

Raffray's bandicoot

Drawing of bandicoot

P. raffrayana
(Milne-Edwards, 1878)

New Guinea
Map of range
Size: 17–39 cm (7–15 in) long, plus 11–23 cm (4–9 in) tail[39]

Habitat: Forest, shrubland, and grassland[41]

Diet: Insects, as well as other invertebrates, small vertebrates, and plants[42]
 LC 


Unknown Template:Population change unknown[41]

Thylacomyidae

Genus MacrotisReid, 1837 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Greater bilby

Gray bilby

M. lagotis
Reid, 1837
Western and central Australia
Map of range
Size: 29–55 cm (11–22 in) long, plus 20–29 cm (8–11 in) tail[43]

Habitat: Savanna and grassland[44]

Diet: Seeds, bulbs, invertebrates, fruit, fungi, and lizards, as well as eggs and small mammals[45]
 VU 


9,000 Population declining[44]

Lesser bilbyExtinct

Brown bilby

M. leucura
Thomas, 1887
Central Australia
Map of range
Size: 20–27 cm (8–11 in) long, plus 12–17 cm (5–7 in) tail[43]

Habitat: Savanna, shrubland, and grassland[46]

Diet: Insects, fruit, seeds, and fungi[47]
 EX 


0 Population steady[46]

Notes

  1. Although it is not recognized as such by the IUCN, Chaeropus ecaudatus is sometimes split into the southern pig-footed bandicoot (C. ecaudatus) and northern pig-footed bandicoot (C. yirratji).

References

  1. "Fossilworks: Peramelemorphia". Paleobiology Database. University of Wisconsin–Madison. http://www.fossilworks.org/cgi-bin/bridge.pl?a=taxonInfo&taxon_no=234407. 
  2. Meredith, R. W.; Westerman, M.; Springer, M. S. (2008). "A timescale and phylogeny for "Bandicoots" (Peramelemorphia: Marsupialia) based on the sequences for five nuclear genes". Molecular Phylogenetics and Evolution 47 (1): 1–20. doi:10.1016/j.ympev.2008.01.002. PMID 18328736. 
  3. Wilson, Reeder, pp. 38–42
  4. 4.0 4.1 Nowak 2005, p. 124
  5. 5.0 5.1 Burbidge, A. A.; Woinarski, J. (2016). "Chaeropus ecaudatus". IUCN Red List of Threatened Species 2016: e.T4322A21965168. doi:10.2305/IUCN.UK.2016-2.RLTS.T4322A21965168.en. https://www.iucnredlist.org/species/4322/21965168. 
  6. 6.0 6.1 6.2 6.3 6.4 6.5 Nowak 2005, p. 128
  7. 7.0 7.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Salas, L.; Seri, L. et al. (2016). "Echymipera clara". IUCN Red List of Threatened Species 2016: e.T7015A21966942. doi:10.2305/IUCN.UK.2016-2.RLTS.T7015A21966942.en. https://www.iucnredlist.org/species/7015/21966942. 
  8. 8.0 8.1 Tran, Nghi (2023). "Echymipera kalubu". University of Michigan. https://animaldiversity.org/accounts/Echymipera_kalubu/. 
  9. 9.0 9.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Helgen, K.; Seri, L. et al. (2016). "Echymipera kalubu". IUCN Red List of Threatened Species 2016: e.T7018A21966845. doi:10.2305/IUCN.UK.2016-2.RLTS.T7018A21966845.en. https://www.iucnredlist.org/species/7018/21966845. 
  10. 10.0 10.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Helgen, K.; Seri, L. (2016). "Echymipera davidi". IUCN Red List of Threatened Species 2016: e.T7017A21966774. doi:10.2305/IUCN.UK.2016-2.RLTS.T7017A21966774.en. https://www.iucnredlist.org/species/7017/21966774. 
  11. 11.0 11.1 Hart, Myha (2012). "Echymipera rufescens". University of Michigan. https://animaldiversity.org/accounts/Echymipera_rufescens/. 
  12. 12.0 12.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Helgen, K.; Seri, L. et al. (2016). "Echymipera rufescens". IUCN Red List of Threatened Species 2016: e.T7019A21966655. doi:10.2305/IUCN.UK.2016-2.RLTS.T7019A21966655.en. https://www.iucnredlist.org/species/7019/21966655. 
  13. 13.0 13.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Helgen, K.; Seri, L. et al. (2016). "Echymipera echinista". IUCN Red List of Threatened Species 2016: e.T7016A21967022. doi:10.2305/IUCN.UK.2016-2.RLTS.T7016A21967022.en. https://www.iucnredlist.org/species/7016/21967022. 
  14. Helgen, K. M.; Flannery, T. F. (2004). "A new species of bandicoot, Microperoryctes aplini, from western New Guinea". Journal of Zoology 264: 117–124. doi:10.1017/S0952836904005667. 
  15. 15.0 15.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F. J.; Helgen, K.; Seri, L. (2019). "Microperoryctes aplini". IUCN Red List of Threatened Species 2019: e.T136538A21965745. doi:10.2305/IUCN.UK.2019-1.RLTS.T136538A21965745.en. https://www.iucnredlist.org/species/136538/21965745. 
  16. 16.0 16.1 16.2 16.3 Nowak 2018, p. 214
  17. 17.0 17.1 17.2 Nowak 2005, p. 130
  18. 18.0 18.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F. J.; Helgen, K.; Seri, L. et al. (2019). "Microperoryctes murina". IUCN Red List of Threatened Species 2019: e.T13389A21965585. doi:10.2305/IUCN.UK.2019-1.RLTS.T13389A21965585.en. https://www.iucnredlist.org/species/13389/21965585. 
  19. 19.0 19.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Helgen, K.; Seri, L. et al. (2016). "Microperoryctes papuensis". IUCN Red List of Threatened Species 2016: e.T13390A21965507. doi:10.2305/IUCN.UK.2016-2.RLTS.T13390A21965507.en. https://www.iucnredlist.org/species/13390/21965507. 
  20. 20.0 20.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F. J.; Helgen, K.; Seri, L. et al. (2016). "Microperoryctes longicauda". IUCN Red List of Threatened Species 2016: e.T84783217A21965649. doi:10.2305/IUCN.UK.2016-3.RLTS.T84783217A21965649.en. https://www.iucnredlist.org/species/84783217/21965649. 
  21. 21.0 21.1 Nowak 2005, p. 129
  22. 22.0 22.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Helgen, K.; Seri, L. et al. (2016). "Rhynchomeles prattorum". IUCN Red List of Threatened Species 2016: e.T19711A21967091. doi:10.2305/IUCN.UK.2016-2.RLTS.T19711A21967091.en. https://www.iucnredlist.org/species/19711/21967091. 
  23. Strahan, p. 173
  24. 24.0 24.1 Burbidge, A. A.; Woinarski, J. (2016). "Isoodon auratus". IUCN Red List of Threatened Species 2016: e.T10863A115100163. doi:10.2305/IUCN.UK.2016-3.RLTS.T10863A21966258.en. https://www.iucnredlist.org/species/10863/115100163. {{cite iucn}}: error: |doi= / |page= mismatch (help)
  25. Norlin, Molly (2012). "Isoodon auratus". University of Michigan. https://animaldiversity.org/accounts/Isoodon_auratus/. 
  26. 26.0 26.1 Szaura, Troy (2018). "Isoodon macrourus". University of Michigan. https://animaldiversity.org/accounts/Isoodon_macrourus/. 
  27. 27.0 27.1 Lunney, D.; Dickman, C.; Woinarski, J. (2016). "Isoodon macrourus". IUCN Red List of Threatened Species 2016: e.T40552A21966494. doi:10.2305/IUCN.UK.2016-2.RLTS.T40552A21966494.en. https://www.iucnredlist.org/species/40552/21966494. 
  28. Strahan, p. 176
  29. 29.0 29.1 Burbidge, A. A.; Woinarski, J. (2016). "Isoodon obesulus". IUCN Red List of Threatened Species 2016: e.T40553A115173603. doi:10.2305/IUCN.UK.2016-3.RLTS.T40553A21966368.en. https://www.iucnredlist.org/species/40553/115173603. {{cite iucn}}: error: |doi= / |page= mismatch (help)
  30. Blevins, Kasey (2022). "Isoodon obesulus". University of Michigan. https://animaldiversity.org/accounts/Isoodon_obesulus/. 
  31. 31.0 31.1 31.2 31.3 31.4 Nowak 2005, p. 126
  32. 32.0 32.1 Burbidge, A. A.; Woinarski, J. (2016). "Perameles eremiana". IUCN Red List of Threatened Species 2016: e.T16570A21965953. doi:10.2305/IUCN.UK.2016-3.RLTS.T16570A21965953.en. https://www.iucnredlist.org/species/16570/21965953. 
  33. 33.0 33.1 Woinarski, J.; Burbidge, A. A. (2016). "Perameles gunnii". IUCN Red List of Threatened Species 2016: e.T16572A21966027. doi:10.2305/IUCN.UK.2016-2.RLTS.T16572A21966027.en. https://www.iucnredlist.org/species/16572/21966027. 
  34. Lancaster, Eric (2001). "Perameles gunnii". University of Michigan. https://animaldiversity.org/accounts/Perameles_gunnii/. 
  35. 35.0 35.1 Lunney, D.; Dickman, C.; Menkhorst, P. (2016). "Perameles nasuta". IUCN Red List of Threatened Species 2016: e.T40554A115173969. doi:10.2305/IUCN.UK.2016-3.RLTS.T40554A21966169.en. https://www.iucnredlist.org/species/40554/115173969. {{cite iucn}}: error: |doi= / |page= mismatch (help)
  36. Adam, Ryan (2015). "Perameles nasuta". University of Michigan. https://animaldiversity.org/accounts/Perameles_nasuta/. 
  37. 37.0 37.1 Burbidge, A. A.; Woinarski, J. (2016). "Perameles bougainville". IUCN Red List of Threatened Species 2016: e.T16569A21965819. doi:10.2305/IUCN.UK.2016-1.RLTS.T16569A21965819.en. https://www.iucnredlist.org/species/16569/21965819. 
  38. Rochefort, Laura (2013). "Perameles bougainville". University of Michigan. https://animaldiversity.org/accounts/Perameles_bougainville/. 
  39. 39.0 39.1 Nowak 2018, p. 208
  40. 40.0 40.1 40.2 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Helgen, K.; Seri, L. et al. (2016). "Peroryctes broadbenti". IUCN Red List of Threatened Species 2016: e.T16710A21965270. doi:10.2305/IUCN.UK.2016-2.RLTS.T16710A21965270.en. https://www.iucnredlist.org/species/16710/21965270. 
  41. 41.0 41.1 Leary, T.; Wright, D.; Hamilton, S.; Singadan, R.; Menzies, J.; Bonaccorso, F.; Helgen, K.; Seri, L. et al. (2016). "Peroryctes raffrayana". IUCN Red List of Threatened Species 2016: e.T16711A21965412. doi:10.2305/IUCN.UK.2016-2.RLTS.T16711A21965412.en. https://www.iucnredlist.org/species/16711/21965412. 
  42. Kennedy, Karen (2023). "Peroryctes raffrayana". University of Michigan. https://animaldiversity.org/accounts/Peroryctes_raffrayana/. 
  43. 43.0 43.1 Nowak 2005, p. 122
  44. 44.0 44.1 Burbidge, A. A.; Woinarski, J. (2016). "Macrotis lagotis". IUCN Red List of Threatened Species 2016: e.T12650A21967189. doi:10.2305/IUCN.UK.2016-2.RLTS.T12650A21967189.en. https://www.iucnredlist.org/species/12650/21967189. 
  45. Brown, Emily (2011). "Macrotis lagotis". University of Michigan. https://animaldiversity.org/accounts/Macrotis_lagotis/. 
  46. 46.0 46.1 Burbidge, A. A.; Woinarski, J. (2016). "Macrotis leucura". IUCN Red List of Threatened Species 2016: e.T12651A21967376. doi:10.2305/IUCN.UK.2016-2.RLTS.T12651A21967376.en. https://www.iucnredlist.org/species/12651/21967376. 
  47. Singh, Angela (2001). "Macrotis leucura". University of Michigan. https://animaldiversity.org/accounts/Macrotis_leucura/. 

Sources

  • Mammal Species of the World. 1 (3rd ed.). Johns Hopkins University Press. 2005. ISBN 978-0-8018-8221-0. 
  • Nowak, Ronald M. (2005). Walker's Marsupials of the World. Johns Hopkins University Press. ISBN 978-0-8018-8222-7. 
  • Nowak, Ronald M. (2018). Walker's Mammals of the World. Monotremes, Marsupials, Afrotherians, Xenarthrans, and Sundatherians. Johns Hopkins University Press. ISBN 978-1-4214-2468-2. 
  • Strahan, Ronald (1995). Mammals of Australia. Smithsonian Institution. ISBN 978-1-56098-673-7.